Description. Frontoparietals: Fragmentarily preserved remains are covered dorsally with the characteristic pit-and-ridge style sculpture as well as low spines (Fig. 7 A, H). The tectum supraorbitale is moderately broad. In dorsal view, the lateral superior process is longer than broad. The articulation surface with the squamosal is well-developed, concave and oriented fully laterally (Fig. 7 A). Neither incrassation frontoparietalis nor the margins are preserved / observable in the remains. The paroccipital process is reduced, but its dorsal surface possesses a distinct crest. The medial base of the paroccipital process is pierced by the occipital arterial foramen, which is covered dorsally by the posterior margin of the frontoparietal and, thus, not visible in dorsal view (Fig. 7 E). The fragmentarily preserved parts of the inferior superior process suggest that it was not longer than the lateral superior process. However, its lateral and slightly ventrally bending can be assumed. The arteria orbitonasalis opens ventrally on the supraorbital tectum (Fig. 7 F). Squamosals: The dorsal surface is covered by a similar to frontoparietal pit-and-ridge sculpture. The bone remains are fragmentarily preserved. Only in HLMD-Ez 2105 (Fig. 7 H, I), an intact posterodorsal process lamina is present, and shows rounded posterior margin. The dorsal and zygomatic processes are broken. However, considering the presence of intact and concave bone margins between the preserved bases of these processes, we can suggest that they were delimited from each other. In ventral view, lamellar structures at the base of the posterolateral processes are observable (Fig. 7 I, K). However, they are incomplete for any further description. Maxillae: The labial surface of the bone is covered by a dense network of moderately deep to deep pits-and-ridge sculpture. The distinct zygomatic process extends posterodorsally and has a rounded posterior tip (Fig. 5 L). The posterior process projects backwards. It is separated clearly from the zygomatic process by a deep concavity and projects much posteriorly. The orbital margin is concave. In lingual view, the pterygoid process projects posteromedially (Fig. 7 M, O). Anteriorly, it is connected with the dorsally oriented lamella above the horizontal lamina. Premaxilla: In anterior view, the pars dentalis is low but broad. Its surface is covered by rugose structures (Fig. 7 P), which recalls the pit-and-ridge sculpture of, e. g., frontoparietal and maxilla. The pars facialis is broken, but its preserved base suggests a L-shaped form. Medially from this process, another shorter and posteromedially oriented process is present. The lateral crest (sensu Venczel 2004) is moderately developed. The dental crest possesses 15 tooth pedicles (Fig. 7 Q). Vertebrae: three presacral and two sacral vertebrae are present. The vertebra centrum is procoelous (four vertebrae, HLMD-Ez 2098, 2100 - 2102) or amphicoelous (one vertebra, HLMD-Ez 2099, Fig. 7 R). The HLMD-Ez 2099 represents a small-sized individual that, most probably, does not have fully ossified joints. In presacral vertebrae, the neural arch is high; and the neural canal is large. The foramina for the spinal nerve are present slightly above the posterior bases of the neural arch. In presacral vertebrae, they are visible in lateral view, whereas in sacral ones - in posterior views (Fig. 7 T). The sacral vertebrae have a broad and flattened transverse process. Ilium: Though all ilia are very fragmentarily preserved, the following characters can be observed on the material: the acetabular region triangular; the acetabulum itself has round outline; the dorsal prominence low and covered by rare irregular structures; the dorsal protuberance absent; the moderately deep spiral groove extends from ventrolateral to dorsomedial direction in the region of the fusion of iliac shaft and acetabulum (Fig. 7 W - AA); in medial view, the ilioischiatic juncture is covered by elongated striae (Fig. 7 Z); the dorsal acetabular expansion larger than the ventral one.
Vasilyan, Davit, Cernansky, Andrej, Szyndlar, Zbigniew, Moers, Thomas (2022): Amphibian and reptilian fauna from the early Miocene of Echzell, Germany. Fossil Record 25 (1): 99-145, DOI: http://dx.doi.org/10.3897/fr.25.83781, URL: http://dx.doi.org/10.3897/fr.25.83781