Hyperolius bocagei

Material (74 specimens, 10 tadpole lots): PEM A 12423, Cunde waterfall, - 13.77390 ° 18.75520 °, 1,287 m asl; PEM A 12455 – 60, PEM A 14085 – 6 (tadpoles) Cuanavale River source lake, - 13.08537 ° 18.89098 °, 1,340 m asl; PEM A 12489, PEM A 14098 (tadpoles), INBAC (no number), Dala River, near Samanunga village, - 12.93169 ° 18.81458 °, 1,363 m asl; PEM A 12508 – 12, river crossing before Samboano village, - 12.30700 ° 18.62350 °, 1,397 m asl; PEM A 12543 – 52, INBAC (no number x 2), Cuito River source lake, - 12.68935 ° 18.36012 °, 1,431 m asl; PEM A 12592 – 8, Calua River source lake, 6 km SE of Cuito River source, - 12.73675 ° 18.39310 °, 1,446 m asl; PEM A 12619, Stop 1: road to Cuito River source, - 12.25050 ° 18.63730 °, 1,556 m asl; PEM A 12674, Camp 3, Malova Village, Mipanha River, - 14.09140 ° 16.41476 °, 1,553 m asl; PEM A 12734 – 7, INBAC: WC- 4833, INBAC (no number x 2), Cuando River source, - 13.00346 ° 19.12751 °, 1,553 m asl; PEM A 12788 – 9, INBAC: WC- 4614, Quembo River, source trap 2, - 13.13544 ° 19.04397 °, 1,374 m asl; PEM A 14112 (tadpoles), Cuiva River, - 11.98346 ° 17.72841 °, 1,264 m asl; PEM A 12815, PEM A 14125 (tadpoles), Cuiva River source, - 12.66856 ° 18.35307 °, 1,407 m asl; PEM A 12857, INBAC: WC- 4616; INBAC (no number x 5) Quembo River source lake, - 13.13624 ° 19.04591 °, 1,411 m asl; PEM A 13767 – 72, INBAC: WC- 6979 Lungwebungu River, old oxbows, - 12.58129 ° 18.67162 °, 1,304 m asl; PEM A 13800 – 2, PEM A 13803 (tadpoles), Lake Tchanssengwe, - 12.41402 ° 18.64417 °, 1,415 m asl; PEM A 14090 (tadpoles), 4 km upstream from Cuanavale River source, - 13.05084 ° 18.89726 °, 1,394 m asl; PEM A 14103 (tadpoles), 4 km downstream from Cuanavale River source camp, - 13.11585 ° 18.90246 °, 1,354 m asl; PEM A 14106 (tadpoles), confluence of Cuito and Calua rivers, - 13.12458 ° 18.89989 °, 1,345 m asl; PEM A 14686, detour route just across bridge over Cuiva River, - 12.13942 ° 18.39393 °, 1,385 m asl; PEM A 14693 (tadpoles), Lungwebungu River camp, first oxbow on right side, - 12.58117 ° 18.67106 °, 1,294 m asl; PEM A 14717, en route from Samanunga village to Cuanavale River source, - 12.94331 ° 18.81118 °, 1,407 m asl; PEM A 14727 – 8, Quembo River, oxbow near small waterfall, - 13.54257 ° 19.29551 °, 1,233 m asl; PEM A 14748 – 9, PEM A 14763 – 65, PEM A 14778, Quembo River right side tributary (Micongo River) past village, - 13.51877 ° 19.28487 °, 1,248 m asl; PEM A 14757, left side tributary (Condinde River) at Cuando River bridge, - 13.60076 ° 19.52675 ° 1,219 m asl. Additional material (16 tadpole lots): SAIAB 204553 (7 tadpoles), Cuanavale River source lake, - 13.08997 ° 18.89389 °, 1,358 m asl; SAIAB 209059 (15 tadpoles), Cuanavale River near confluence, - 13.12478 ° 18.90017 °, 1,347 m asl; SAIAB 204509 (29 tadpoles), outlet from Kuembo River source lake, - 13.13689 ° 19.03144 °, 1,375 m asl; SAIAB 204564 (25 tadpoles), SAIAB 209074 (1 tadpole), Quembo River source lake, - 13.13611 ° 19.04500 °, 1,366 m asl; SAIAB 209066 (9 tadpoles), Quembo River source lake, - 13.14025 ° 19.04822 °, 1,365 m asl; SAIAB 209062 (2 tadpoles), swamp near Cuanavale River source lake camp, - 13.10750 ° 18.86089 °, 1,386 m asl; SAIAB 204465 (19 tadpoles), Cuando River source pool, - 13.00383 ° 19.12719 °, 1,350 m asl; SAIAB 209060 (2 tadpoles), stream outflow about 2.6 km downstream of source lake, - 13.00317 ° 19.15153 °, 1,331 m asl; SAIAB 204472 (13 tadpoles), Cuando River, - 13.09142 ° 19.35850 °, 1,283 m asl; SAIAB 209067 (8 tadpoles), on Cuanavale River road, - 13.14190 ° 19.44503 °, 1,301 m asl; SAIAB 209070 (10 tadpoles), pan near Cuanavale River camp, seepage, - 13.10750 ° 18.86089 °, 1,387 m asl; SAIAB 204569 (13 tadpoles), Samununga village, - 12.93228 ° 18.81672 °, 1,365 m asl; SAIAB 209063 (6 tadpoles), small stream 3 km below Cuanavale River source camp, - 13.12539 ° 18.89914 °, 1,344 m asl; SAIAB 209069 (11 tadpoles), pool in wetland on road edge, west of Munhango, - 12.17806 ° 18.24306 °, 1,370 m asl; SAIAB 209075 (18 tadpoles), Cuando River source bog, - 13.00383 ° 19.12719 °, 1,350 m asl. Description: Medium sized reed frog; rounded snout; reduced webbing; ventrum granular. Three different color forms were observed, i. e., plain light to dark green, brown, and bright red; thin dorsolateral white stripe present; small white specks on dorsal side of legs; toes and webbing red. Tadpoles with ventrum spotted and ventral blotches remain present in juveniles but fade in adults. Adult females (n = 11) varied from 16.7 – 21.9 (19.6) mm (largest female: PEM A 12549); adult males (n = 52) varied from 14.8 – 21.1 (18.7) mm (largest male: PEM A 12592). Habitat and natural history notes: Found at the margins of large rivers and source lakes within dense emergent wetland vegetation. Sympatric with other species of Hyperolius, mostly H. nasutus and H. parallelus. Comments: Initially we assigned these specimens to H. bocagei (NGOWP 2017), however it is generally regarded as a larger species that is associated with the viridiflavus group (see Schiøtz 1999). This new material differs in size (smaller) and webbing (less webbing) from H. bocagei (Schiøtz 1999). Phylogenetic work is currently underway to determine its taxonomic status.