Description. Adult medusae with smooth exumbrella, with evenly thick mesoglea of rigid consistency (Fig. 3, Fig. 4, Fig. 5). BH about 35 mm and DBW about 50 mm (Table 3). Interradius thickened throughout bell height, with deep median furrow spanning height of bell. Adradial furrows spanning lower two thirds of bell. Coronal indentation shallow just below apex. Exumbrella lacking nematocyst warts or freckles. Gonads attached along entire length of interradial septa, leaf-shaped, not overlapping along the interradius (Fig. 6 A, 7 A). Manubrium length about 40 % of bell height. Manubrium cruciform with four narrow, lanceolate lips (Fig. 6 B). Gastric saccules unbranched, slightly raised and opaque (Fig. 5 D, 6 C, 7 B). Gastric phacellae horseshoe-shaped in each corner of stomach (Fig. 6 D, 7 C). Gastric cirri simple and unbranched (Fig. 6 E, 7 D). Sensory niches four, perradial, U-shaped, with one shallow covering scale above and no lower scale, upper scale with central tongue-shaped flap partially shielding rhopalium, located approximately 1 / 6 of bell height from velarial turnover (Fig. 6 F, G, 7 E, F). The rhopaliar niche located in a triangular shaped depression of exumbrella (Fig. 6 F, 7 E). Subumbrellar rhopalial windows flat (Fig. 6 H). Each of the four rhopalia bearing a set of six eyes, with the two median possessing prominent lenses and the four lateral ones adjacent to the lens eyes being pigment pits and slit (Fig. 6 I-K, Fig. 7 G, H). A single broad bean shaped statolith located behind each complex eye (Fig. 6 I-K). Velarial canals one per octant, dendritic, with only the tips defined along the velarial margin (Fig. 6 L, Fig. 7 I). Frenulum a single solid, gelatinous struc ture, extending to near velarial margin (Fig. 6 L). Velarial width about 20 % of DSW. Pedalia scalpel-like, each bearing one tentacle (Fig. 6 M, Fig. 7 J). Pedalia about half of bell height, inner keel rounded, about two times the width of pedalial canal. Outer keel width approximately twice that of pedalial canal, inner keel width and outer keel width almost equal. Proximal pedalial canal bend slight volcano-shaped (Fig. 6 N, Fig. 7 K). Tentacles four, with one per pedalium, base width up to 2.7 mm thick, round in cross section, with nematocyst rings alternate thick and thin in preserved specimens (Fig. 6 O, 7 L). Color and length of tentacles in living specimens was light brownish (Fig. 3 A, B). The smallest young medusa (NSMT-Co 1570) had a BH of about 15 mm, DBW of about 20 mm (Fig. 8, 9). Mesoglea was thin and softer than that of adults. Adradial furrows spanning half of the bell height (Fig. 8 A, 9 A). Interradial furrows spanning the entire height of the bell (Fig. 8 B, 9 B). Coronal indentation shallow and exumbrella nematocyst freckles absent. Gonads attached along entire length of interradial septa, leaf-shaped, not overlapping along the interradius (Fig. 10 A). Manubrium length about 20 % of bell height. Gastric saccules not developed. Gastric phacellae four-leaf clover shaped in each corner of stomach (Figs 8 C, 9 C, 10 B). Depression of exumbrella not developed. Upper medial rhopaliar scale flap shorter than in adults (Fig. 10 C, D). Velarial canals one per octant, dendritic but with minor branching (Fig. 10 E). Pedalium scalpel-like (Fig. 10 F). Volcano-shaped pedalial canal bend smoother than in adults (Fig. 10 G). Tentacle with nematocyst rings alternate thick and thin in preserved specimens (Fig. 10 H). Cnidome. Six different nematocyst types identified and measured in the paratype specimen (NSMT-Co 1571) (Table 4, Fig. 11). Tentacle: Large microbasic p-rhopaloids, banana-shaped p-mastigophores, small oval beehive isorhizas, rod-shaped isorhizas, small sub-spherical p-rhopaloids. Manubrium: Tiny microbasic euryteles, small sub-spherical p-rhopaloids. Phacellae: Tiny microbasic euryteles, small sub-spherical p-rhopaloids. Exumbrella lacking nematocysts.
Meteoronakishinouyei, a new family, genus and species (Cnidaria, Cubozoa, Chirodropida) from Japanese Waters